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Indications of how this may have come about are afforded by the lower group of the Anacrogynous Jungermanniaceae, and throughout the Acrogynous Jungermanniacae the plant has well-marked stem and leaves.

Lastly Haplomitrium, a rare British genus, forms with the exotic Calobryum, an isolated group which is most naturally placed among the anacrogynous forms although the archegonia are in terminal groups.