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Animal rights activists have recently posted drone footage appearing to show dolphins swimming in murky green tanks on the premises.

For months managers at Marineland have tried to send the killer whales to other marine zoos but this has angered animal rights campaigners who want them housed in a sanctuary, where the orcas won't have to perform or be used for breeding.

Other animal rights groups have been more welcoming of Pannier-Runacher's announcement, saying a European sanctuary would still be better for the orcas' welfare than life in another marine zoo.

The Greyhound Board of Great Britain accused ministers of giving way to pressure from "the extreme animal rights movement".

But the Greyhound Board of Great Britain, which regulates the sport, said the announcement had "nothing to do with greyhound welfare and everything to do with pressure from the extreme animal rights movement".