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The podobranchiae are clearly epipodites, or, more correctly, parts of the epipodites, and it is probable that the arthro- and pleurobranchiae are also epipodial in origin and have migrated from the proximal segment of the limbs on to the adjacent body-wall.

Arthro�dia, a species of articulation, in which the head of one bone is received into a shallow socket in another; a ball-and-socket joint.

The Dissennith also, and the Arthro, flow through Merionethshire and the land of Conan.