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Those include the dwarf cuscus, a large-eyed marsupial; the anoa, a midget buffalo; the Sulawesi crested black macaque, locally known as yaki; and the babirusa, or deer-pig, which is renowned for its large tusks.

The researchers who announced the recent discovery previously described a 35,000-year-old cave painting of a babirusa, or pig deer, there.

For the babirusa, the most recent common ancestor for all specimens was found to be at 2.49m years ago.

The study focused on 14 cave paintings: 12 human hand stencils and two naturalistic animal depictions, one showing an animal called a babirusa, or "pig-deer," and the other showing what probably is a pig.

A painting of an animal known as a pig deer, of the species babirusa, was determined to be at least 35,400 years old.