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Archegonium with a slender persistent style, solitary on a usually very short branch; the perianth free from the involucral leaves, oval or oblong, terete or angular, variously carinate, cristate, or ciliate.

Cones from 4 to 6 cm. long, ovate or ovate-conic, symmetrical; apophyses nut-brown, flat or convex and transversely carinate, the prickle of the umbo more or less persistent.

It must have been that of the struthious birds or that of the carinate birds, or something different from both.

The Rostro– carinate flints found at the base of the Crag are long bars with a beak–end, suited for breaking up earth.

Empty glumes persistent, membranaceous and shining, carinate, acute, nearly equal; flowering glumes toothed or erose-denticulate at the truncate summit, usually delicately 3–5-nerved, with a slender twisted awn near or below the middle.