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Accumbent cotyledons have their edges placed against the caulicle.

The stem at the very beginning produces roots, in most plants a single root from the base of the embryo-stem, or caulicle.

In germination this store is promptly utilized in the development of the caulicle to twenty or thirty times its length in the seed, and to corresponding thickness, in the formation of a cluster of roots at its lower end, and the early production of the incipient plumule; also in their own growth into efficient green leaves.

For they are thickened to the utmost, even into hemispheres; the caulicle does not lengthen at all; merely sends out roots from the lower end, and develops its strong plumule from the upper, the seed remaining unmoved underground.

Embryo of Pea, i. e. a pea with the coats removed; the short and thick caulicle presented to view.