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These “coadapted gene complexes” are broken up with excessive outbreeding.

We conclude that in this animal complex, the gut microbiome and host genome represent a coadapted “hologenome” that breaks down during hybridization, promoting hybrid lethality and assisting speciation.

Coadapted: formed so as to work together to one end; as the mandible and maxilla in Chrysopids, etc.

In order that an animal should acquire some structure specially and largely developed, it is almost indispensable that several other parts should be modified and coadapted.