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"Male bonobos that are more aggressive obtain more copulations with females, which is something that we would not expect," said Mouginot.

Her remembrance of scenes of copulations past causes her no shame.

While males are aggressive with each other, they aren’t aggressive toward females and do not force copulations, despite being twice the size of their mating partners.

Ducks' evolution of a penis is thought to have facilitated males being able to force copulations to gain control of fertilization.

The highly promiscuous ruddy ducks never form pair bonds, and almost all copulations are forced.