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Other safaris, both on foot and in vehicles, focus on the dawn chorus, bats and moths, purple emperor butterflies, Sussex oaks, owls, kingfishers and other highlights of the natural world.

Species like purple emperor butterflies and nightingales, which had not been sighted for a generation, returned, the soil recovered from decades of fertilizer overuse, and natural river systems were restored.

The Knepp Castle estate boasts more purple emperor butterflies than anywhere in Britain.

The results have been astonishing: We now have 2 percent of the U.K.’s population of nightingales and have become a breeding hot spot for numerous nationally rare species, including purple emperor butterflies and turtledoves.

The hackberry tree also feeds the hackberry emperor butterfly, the tawny emperor and the odd-looking snout butterfly.