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The parasite even participates in the equatorial plate that defines metaphase, along with the chromosomes!

Both Theodor Boveri and van Beneden, in their papers of 1887, regarded the centrosome as initiating, not only the division of the cell-body but that of the chromatin also; Beneden even suggested that the pull of the mantle fibres caused the division of the chromatin in the equatorial plate.

W. Pfitzner in 1882 was the first to show that the splitting of the chromosomes in the equatorial plate was only the reappearance of a split in the spireme thread and was due to a corresponding 715 division into two of each of the chromatin granules.

The metaphase is the parting of the sister chromosomes in the equatorial plate; their passage to opposite poles of the spindle constitutes the anaphase; and their reconstruction to form the resting daughter nuclei, the telophase.

Equatorial plate of spermatogonium, 30 chromosomes—29 large, 1 small.