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We identified three major loci and two minor loci responsible for exserted stigma, and found that all the five QTLs were located within domestication sweeps.

Palpi exserted, approximating, covered with scales, but without hairs, the last joint naked, slender, acute.

Erect culms and appressed leaves more slender than in the preceding; panicle exserted, very simple and narrow; spikelets smaller, the lower glumes acuminate, little shorter than the cuspidate upper one.

Included, inclosed by the surrounding organs; not exserted.

They are hypogynous, and have long and very delicate filaments, and large, linear or oblong two-celled anthers, dorsifixed and ultimately very versatile, deeply indented at each end, and commonly exserted and pendulous.