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Dr Shuttleworth said "we don't need a grey squirrel to set foot on the island for pox to come here" as reds moved between Anglesey and Gwynedd on the mainland.

During the fourth annual survey week in October, 659 participants reported 255 grey squirrel and 510 red squirrel sightings - more than triple the number reported in a typical week.

The traditional way of managing the grey squirrel population is by culling them.

A plan to use oral contraceptives to control grey squirrel populations in the UK is making good progress and could soon be put to the test in field trials, say government scientists.

The project could help eradicate the grey squirrel in the UK without killing them, says environment minister Lord Benyon.