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The chief distinctive characters of the sporogenous hyphae are their orientation, usually vertical; their limited apical growth; their peculiar branching, form, colour, contents, consistency; and their spore-production.

The series of increasingly complex sporogonia among Bryophytes appears to be most naturally explained on an hypothesis of progressive sterilization of sporogenous tissue, such as has been advanced by Bower.

Other series of modifications arise in which the tissues corresponding to the stroma invest the sporogenous hyphal ends, and thus enclose the spores, asci, basidia, &c., in a cavity.

The cells which will produce the sporogenous tissue are shaded.