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As the one wave ascends while the other descends, a pressure of the basilar membrane occurs at the point where they meet; this causes the basilar membrane to move towards the tectorial membrane, forcing this membrane suddenly against the apices of the hair-cells, thus irritating the nerves.

The hairs of the sense cells are shaken by the vibration of the water, and pass the impulse back to the end-brushes of the auditory axons, The tectorial membrane looks as if it might act as a damper, but may be concerned, as "accessory apparatus," in the stimulation of the hair cells.