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Each cone consists of an axis, on which numerous broad and thin bracts are arranged in regular rows; in the axil of each bract occurs a single flower; a male flower is enclosed by two opposite pairs of leaves, forming a perianth surrounding a central sterile ovule encircled by a ring of stamens united below, but free distally as short filaments, each of which terminates in a trilocular anther.

In the coco-nut, a trilocular and triovular ovary produces a one-celled, one-seeded fruit.

Trilocular, three-celled, as the pistils or pods in fig. 328-330.

In the Pucciniæi the distinctive features of the genera are based upon the more or less complex nature of the pseudospores, which are bilocular in Puccinia, trilocular in Triphragmium, multilocular in Phragmidium, &c.