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A necropsy report also revealed that one of the couple's three dogs, which had been crated while recovering from surgery, died from starvation and dehydration.

Her 14-ton carcass offers several immediate clues, said Keith Matassa, chief executive of the Ocean Animal Response and Research Alliance, who is leading a necropsy of the whale to determine her cause of death.

According to necropsies performed on recovered whales, many of them died from collisions with ships.

Officials performed a necropsy, an autopsy for animals, and tissue samples were taken to be analysed.

The lion weighed nearly 53 pounds and had nothing in her stomach during the attack, according to the necropsy results released from the California Department of Fish and Wildlife last week.